In the third season premiere, Jane and Maura'’s friendship suffers following the shooting of Maura'’s biological father, gangster Paddy Doyle (guest star JOHN DOMAN -– The Wire, Borgia). Jane also finds herself in the middle of an Internal Affairs investigation —and her relationship with Agent Dean (recurring guest star BILLY BURKE -– Twilight films) is exposed. Recurring guest stars JACQUELINE BISSET (Nip/Tuck, Joan of Arc) and ED BEGLEY JR. (St. Elsewhere, Pineapple Express) also appear.